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Pakistan | Family List | Pakistan V. 207

Asteraceae tribe Gnaphalieae


Department of Botany, University of Karachi, Karachi-75270, Pakistan

Herbs or small shrubs, mostly woolly tomentose. Leaves alternate or opposite, sessile or stalked with flat, involute or revolute margins. Capitula solitary or in groups, disciform, discoid rarely radiate; phyllaries 1-multiseriate, entirely scarious, hyaline, white, pink, purple or yellow to herbaceous with scarious margins, stereome divided or undivided. Receptacle flat or ± convex, mostly epaleate, rarely paleate. Outer florets often missing, usually filiform, or radiate when present, bisexual in 1-many rows, usually outnumbering the disc florets, disc florets either perfect or functionally male. Anthers ecalcarate, mostly caudate, rarely ecaudate (Phagnalon), apical appendages often demarcated from the connective, ovate, sometimes oblong, cordate or concave. Style usually with oblong-linear, truncate hairs restricted to upper portion of branch and very often concentrated at the apices of the branches, rarely on the shaft below the branches, hairs obtuse, stigmatic areas in two separate lines. Cypselas small, hairy, sometimes papillose, oblong to obovoid; mostly pappose, rarely epappose pappus mostly monomorphic made up of capillary setae, setae barbellate, rarely plumose; sometimes dimorphic with scales and setae.

A large tribe with 182 genera and more than 2000 species, cosmopolitan in distribution but mostly in southern hemisphere such as southern Australia and Africa. Represented in Pakistan by 12 genera and 45 species.

1 Capitula radiate. Pappus apically plumose.   Leysera
+ Capitula disciform. Pappus mostly barbellate or absent, if apically plumose capitula always disciform.   (2)
2 (1) Cypselas of outer florets (female) epappose and of inner florets pappose; pappus deciduous or the inner florets also epappose. Phyllaries chartaceous or apically papillose.   (3)
+ All cypselas pappose. Phyllaries not as above.   (4)
3 (2) Cypselas and female corollas completely enclosed within phyllaries and always dispersed within them. Corolla obliquely attached to the cypselas.   Cymbolaena
+ Cypselas and female corollas not enclosed within phyllaries, cypselas sometimes included within phyllaries but free and not dispersed within them. Corolla not obliquely attached to the cypselas.   Filago
4 (2) Pappus setae apically plumose.   (5)
+ Pappus setae barbellate.   (6)
5 (4) Capitula few together in globose glomerules. Disc florets purple.   Lasiopogon
+ Capitula few together along the axis forming the leafy spike. Disc florets yellow.   Ifloga
6 (4) Capitula crowded into head, surrounded by the inflorescence bracts, different in texture of indumentum and colour from the rest of the cauline leaves forming a distinct star-like structure; sometimes neither prominent star formed nor the inflorescence bracts different from the other cauline leaves than the cypselas c. 1 mm long; pappus setae (6-) 7-9 times as long as the cypselas (L. nanum).   Leontopodium
+ Inflorescence of 1 to many capitula, sometimes crowded into a head, then not surrounded by radiating star-shaped bracts. Cypselas usually longer; pappus setae 3-4 (-5) times longer than the cypselas.   (7)
7 (6) Anthers completely ecaudate with obtuse bases (in our species)   Phagnalon
+ Anthers either caudate or with sagitatte bases.   (8)
8 (7) Phyllaries white, rarely pinkish, opaque.   (9)
+ Phyllaries brownish or citron to golden yellow.   (10)
9 (8) Capitula dimorphic either female with disc florets male or functionally female or male with a few outer female florets. Cypselas with short clavate not myxogenic hairs.   Anaphalis
+ Capitula not dimorphic, with outer female and disc florets bisexual. Cypselas with short clavate myxogenic hairs.   Pseudognaphalium
10 (8) Pappus setae basally connate and falling as a unit.   Gamochaeta
+ Pappus setae free and not falling as a unit.   (11)
11 (10) Stereome undivided.   Gnaphalium
+ Stereome divided.   (12)
12 (11) Capitula in small loose terminal, flat topped corymbs.   Pseudognaphalium
+ Capitula either few together or densely congested but not in flat topped corymbs.   (13)
13 (12) Capitula densely congested, phyllaries hyaline. All florets yellow.   Homognaphalium
+ Capitula few together. Phyllaries brownish. All florets purple.   Lasiopogon

  • List of lower taxa


    Related Objects  

    Flora of China  
  • Gnaphalieae (PDF)
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