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Pakistan | Family List | Pakistan V. 206 | Cyperaceae | Cyperus

2. Cyperus subg. Pycnostachys C.B. Clarke


1 (1) Robust or sturdy perennials; caespitose or with a creeping, ligneous rhizome (but, first-season plants of C. aucheri resemble annuals, 4).   (1)
+ Short-lived perennials or slender annuals; roots fibrous.   (2)
       
2 (1) Leaf blades reduced; Primary rays and foliose involucral bracts and primary rays up to 20.   22 C. alternifolius
+ Leaf blades well developed; primary rays and involucral bracts up to 5.   (3)
       
3 (2) Caespitose plants with tillers, old dark tillers remain tightly attached to base of new shoots; inflorescence a single head with 4-20 compressed spikes.   28 C. niveus
+ Rhizome long-creeping (mostly deep in sand) giving rise to aerial shoots, or plants caespitose; inflorescence mostly a multiple spike or formed by several capitula, sometimes in stunted plants (notably C. conglomeratus subsp. curvulus) inflorescence reduced to few or single spike.   (4)
       
4 (3) Nuts winged; roots tomentose.   27 C. aucheri
+ Nuts not winged; roots tomentose or glabrous.   (5)
       
5 (5) Caespitose plants or rhizome short; roots tomentose.   (5)
+ Rhizome long-creeping, roots glabrous.   23 C. arenarius
       
6 Plants tall; inflorescence a compact globose head.   26 C. pachyrhizus
+ Plants small; inflorescence an anthelodium.   (7)
       
7 (6) Plants rather small and slender; basal sheaths soft; roots glabrous.   24 C. atkinsoni
+ Plants rigid; basal sheeths hard; roots tomentose.   25 C. conglomeratus
       
8 (8) Plants 10-40 cm; glumes obtuse or acute not recurved.   (8)
+ Plants 4-15 cm; glumes apiculate, conspicuously recurved.   (9)
       
9 (9) Distal partial inflorescence with up to 15 spikes; ligule 0.   (9)
+ Distal partial inflorescence with up to 100 crowded spikes; ligule present.   (10)
       
10 (9) Glumes 1.3-1.5 mm, straight; nut 0.5-0.7 mm, reticulate.   29 C. haspan
+ Glumes 0.7-0.9 mm, rather cymbiform; nut 0.4-0.5 mm, papillose.   30 C. tenuispica
       
11 Spikes in globose heads of up to 100 spikes; glumes distinctly cymbiform, obtuse.   31 C. difformis
+ Spikes in digitate groups of up to 40 spikes; glumes not distinctly cymbiform, acute.   32 C. fuscus
       
12 Glumes truncate, with 3-nerved keel and nerveless sides.   33 C. cuspidatus
+ Glumes attenuate, sides conspicuously nerved.   (13)
       
13 (12) Glumes 2.2-3 mm, sides with 2-3 nerves; stamens 3.   35 C. steadii
+ Glumes 1.5-2.3 mm, sides with 3-4 nerves; stamens 1-2.   34 C. squarrosus


 

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