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BFNA | Family List | BFNA Vol. 3 | Jungermanniaceae

Tritomaria Loeske, Hedwigia. 49: 13. 1909.

Won Shic Hong

Plants ca. 0.5--5 cm ´ 1--4 mm, prostrate to ascending. Stem transverse section with 1--3 cortical cells and 15--25 medullary cells; rhizoids numerous in dense mats, colorless to pale brown. Leaves subtransversely-obliquely inserted, 2--4-lobed, asymmetrical (except T. polita); lobes triangular, unequal, margins entire; cell walls collenchymatous, trigones bulging; oil bodies 3--8 per cell, finely granular, spherical to ovoid; underleaves absent. Sexual condition dioicous. Specialized asexual reproduction by gemmae. Androecia intercalary, antheridial stalk 1--2-seriate, female bracts leaflike, 3--4-lobed, margins entire-dentate, bracteoles usually absent. Perianth cylindrical-ovoid, plicate, mouth entire or ciliate. Capsule oval, walls 3--5-stratose, exterior wall with nodular and interior wall with semiannular bands. Elaters ca. 6--10 m m diam. Spores ca. 10--16 m m.

Species 8 (6 in the flora): North America, Europe, Asia, Africa.


Schuster, R. M. 1969. Tritomaria Schiffn. In: R. M. Schuster. 1966--1993. The Hepaticae and Anthocerotae of North America. New York. Vol. 2, pp. 638--704. Schuster, R. M. 1984. Evolution, Phylogeny and Classification of the Hepaticae. In: R. M. Schuster, ed. 1984. New Manual of Bryology. Vol. 2, pp. 892--1070. Nichinan.


Grolle, R. and D. G. Long. 2000. An annotated check-list of the Hepa;ticae and Anthocerotae of Europe and Macaronesia. J. Bryol. 22: 103--140.
Schuster, R.M. 1958. Notes on Nearctic Hepaticae XIII The genus Tritomaria (Lophoziaceae) in Arctic Canada. Canad. Jour. Bot. 36: 269--288.
Schuster, R. M. 1988. The Hepaticae of South Greenland. Nova Hedwigia. Berlin. Vol. 92.
Schuster, R.M. & K. Damsholt. 1974. The Hepaticae of west Greenland from ca. 60° N to 72° N. Copenhagen.

1 Leaves more or less symmetrical, equally 2--4-lobed with blunt to rounded apices; perianth mouth entire or subentire.   Tritomaria polita
+ Leaves asymmetrical, unequally 2--3-lobed with acute or acuminate apices; perianth mouth dentate.   (2)
2 (1) Leaves complicate to explanate, wider than long; gemmae rare-absent, yellowish-reddish brown.   (3)
+ Leaves canaliculate-complanate, longer than wide; gemmae abundant, reddish brown to rust red.   (4)
3 (2) Plants 1--1.5 mm wide, pellucid; leaves approaching symmetrical form, loosely complex, trigones coarsely nodose; gemmae angulate; perianth mouth teeth 1--2 cells long.   Tritomaria heterophylla
+ Plants 2--3.5 mm wide, opaque; leaves asymmetrical, explanate, trigones moderate-bulging, but never nodose; gemmae generally absent; perianth mouth teeth 4--5 cells long.   Tritomaria quinquedentata
4 (2) Leaves usually 3-lobed, lobes acute to subacute; trigones large; cilia of perianth mouth 1--4 cells long; gemmae reddish-brown.   Tritomaria scitula
+ Leaves asymmetrically 2--3-lobed, lobes acute or acuminate; trigones small; cilia of perianth mouth 5--6 cells long; gemmae rust-red.   (5)
5 (4) Gemmae smooth, ovoid-elliptical; leaf cells 8--14 ´ 10--20 m m in midleaf; cell walls thick, trigones not bulging.   Tritomaria exsecta
+ Gemmae angulate, polygonal; leaf cells 18--22 ´ 20--35 m m in midleaf; cell walls thin, trigones bulging.   Tritomaria exsectiformis

Lower Taxa

Related Synonym(s):


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