1 |
Flowers solitary, axillary; plant dwarf, usually a prostrate shrub; leaf blade small, to 1.1(–1.2) cm. |
|
(2) |
+ |
Inflorescence racemose or corymbose; plant usually a taller shrub; leaf blade at least 1.5 cm. |
|
(10) |
|
|
|
|
2 (1) |
Leaf blade abaxially densely or sparsely setulose. |
|
(3) |
+ |
Leaf blade abaxially glabrous, rarely scattered hispidulous. |
|
(5) |
|
|
|
|
3 (2) |
Flower 4-merous; leaf blade 3–7 × 2–5 mm. |
|
24 Gaultheria suborbicularis |
+ |
Flower 5-merous; leaf blade 0.5–2 cm × 3–10(–16) mm. |
|
(4) |
|
|
|
|
4 (3) |
Leaf blade broadly ovate or suborbicular, sometimes ovate or elliptic, 5–10(–18) × 3–16 mm; corolla
white, pink, or crimson, ca. 5 mm, lobes ca. 1 mm. |
|
25 Gaultheria nummularioides |
+ |
Leaf blade obovate-elliptic or obovate-oblong, 1–2 × 0.5–1 cm; corolla white, 6–9 mm, lobes 3–6 mm. |
|
31 Gaultheria jingdongensis |
|
|
|
|
5 (2) |
Leaves unequal in size; larger leaf blades usually obovate or oblong, less than 2 × as long as wide,
8–14 × 6–8 mm, apex obtuse to rounded. |
|
29 Gaultheria hypochlora |
+ |
Leaves equal, or slightly unequal in size; leaf blade usually elliptic, elliptic-oblong, or linear, more than 2 × as long as wide, 5–16 × 1–7 mm, apex acute, obtuse, or acuminate. |
|
(6) |
|
|
|
|
6 (5) |
Twigs and often leaves glabrous; leaf blade obtuse or acuminate at apex. |
|
32 Gaultheria taiwaniana |
+ |
Twigs and leaves ± hairy; leaf blade obtuse to acute at apex. |
|
(7) |
|
|
|
|
7 (6) |
Pedicel 5–9(–19) mm; leaf blade glabrous. |
|
26 Gaultheria dolichopoda |
+ |
Pedicel 1–2 mm; leaf blade usually variously pubescent or hispidulous, and/or ciliate at margins. |
|
(8) |
|
|
|
|
8 (7) |
Leaf blade long ciliate along margin. |
|
27 Gaultheria trichophylla |
+ |
Leaf blade not long ciliate along margin. |
|
(9) |
|
|
|
|
9 (8) |
Fruit usually white; leaf blade linear or oblong-linear, 5–16 × 1–4 mm; calyx lobes ovate, base
slightly cordate. |
|
28 Gaultheria cardiosepala |
+ |
Fruit usually dark blue; leaf blade elliptic or oblong, 6–16 × 2–6 mm; calyx lobes broadly ovate
or ovate-triangular, base rounded. |
|
30 Gaultheria sinensis |
|
|
|
|
10 (1) |
Leaf blade less than 3.5 cm; plant usually to 40(–60) cm tall. |
|
(11) |
+ |
Leaf blade more than 4 cm; plant usually more than 40 cm tall. |
|
(17) |
|
|
|
|
11 (10) |
Inflorescence racemose, rachis 1–4.5 cm. |
|
(12) |
+ |
Inflorescence shortly racemose, rachis 2–15 mm. |
|
(14) |
|
|
|
|
12 (11) |
Leaves purple abaxially; fruit dark purple, fleshy; leaf blade less than 2 × as long as wide; rachis
glabrous. |
|
22 Gaultheria purpurea |
+ |
Leaves pale green abaxially; fruit whitish, succulent; leaf blade usually more than 2 × as long as wide; rachis densely puberulous. |
|
(13) |
|
|
|
|
13 (12) |
Inflorescence solitary on each twig, pseudoterminal; leaf blade 8–11(–15) × 3–7 mm, elliptic-oblong
or oblanceolate. |
|
6 Gaultheria borneensis |
+ |
Inflorescences usually 2 or 3, terminal and from upper leaf axils; leaf blade 1.2–3.5 × 0.4–1.2 cm,
obovate or narrowly obovate. |
|
9 Gaultheria cuneata |
|
|
|
|
14 (11) |
Plant 30–40 cm tall; twigs rust-colored long setose; inflorescence 3–8-flowered, pedicels ca. 6 mm; leaf
blade ovate, 2–3.5 × 1.5–2 cm. |
|
3 Gaultheria notabilis |
+ |
Plant 5–20 cm tall; twigs pubescent to glabrous; inflorescence 1–5-flowered, pedicel 1–6 mm; leaf blade elliptic, obovate, or suborbicular, 0.8–5 × 0.6–2.5 cm. |
|
(15) |
|
|
|
|
15 (14) |
Calyx and corolla lobes 4; leaf blade hirtellous abaxially. |
|
23 Gaultheria heteromera |
+ |
Calyx and corolla lobes 5; leaf blade punctate or hispidulous abaxially. |
|
(16) |
|
|
|
|
16 (15) |
Corolla campanulate, ca. 4 mm; inflorescence 1–3-flowered; leaf blade elliptic, 10–12 × 6–8 mm, abaxially dark punctate. |
|
7 Gaultheria prostrata |
+ |
Corolla urceolate, ca. 5 mm; inflorescence 2–5-flowered; leaf blade obovate, elliptic, or suborbicular, 13–50 × 8–25 mm, abaxially hispidulous. |
|
8 Gaultheria pyrolifolia |
|
|
|
|
17 (10) |
Twigs glabrous. |
|
(18) |
+ |
Twigs densely hirsute, hispid, or setose. |
|
(24) |
|
|
|
|
18 (17) |
Leaf blade ovate or broadly ovate, base rounded to auriculate-cordate, apex acuminate to caudate-acuminate. |
|
(19) |
+ |
Leaf blade elliptic, oblong, or obovate to lanceolate, base cuneate, sometimes rounded-obtuse, apex acute, sometimes acuminate. |
|
(21) |
|
|
|
|
19 (18) |
Inflorescence corymbose or shortly racemose, axis 3–10 mm; secondary veins 1 pair arising near base
of leaf blade. |
|
2 Gaultheria dumicola |
+ |
Inflorescence racemose, axis at least 3 cm; secondary veins more than 3 pairs. |
|
(20) |
|
|
|
|
20 (19) |
Leaf blade auriculate-cordate at base, abaxially shortly glandular setose, secondary veins 4–7(or 8)
pairs; twigs trigonous and winged; rachis and pedicels pubescent. |
|
11 Gaultheria brevistipes |
+ |
Leaf blade obtuse to subcordate at base, abaxially glabrous, secondary veins 3 or 4 pairs; twigs
terete; rachis and pedicels glabrous. |
|
21 Gaultheria leucocarpa |
|
|
|
|
21 (18) |
Twigs distinctly trigonous or with narrow wings; inflorescence 2–3 cm; bract ovate-lanceolate,
5–6 mm, with 1 distinct main vein, distinctly carinate, secondary veins 2 pairs. |
|
19 Gaultheria trigonoclada |
+ |
Twigs terete, sometimes obtusely angled; inflorescence 2–12 cm; bract ovate to orbicular-ovate, 1–4(–7) mm, main veins inconspicuous, secondary veins more than 3 pairs. |
|
(22) |
|
|
|
|
22 (21) |
Inflorescence terminal and axillary, 10–12 cm; leaf blade obovate or elliptic-obovate, 3–6.5 ×
1–2.5 cm, apex acute, adaxially puberulous on midvein. |
|
16 Gaultheria longiracemosa |
+ |
Inflorescence axillary, 2–9 cm; leaf blade oblong or elliptic, 5–17 × 2–6.5 cm, apex acute to
caudate-acuminate, adaxially glabrous. |
|
(23) |
|
|
|
|
23 (22) |
Leaf blade usually oblong, thickly leathery, apex caudate-acuminate, abaxially densely punctate; bracteoles below middle to near base of pedicel. |
|
18 Gaultheria griffithiana |
+ |
Leaf blade usually elliptic, leathery, apex acute to acuminate, abaxially scattered punctate;
bracteoles close to calyx. |
|
20 Gaultheria fragrantissima |
|
|
|
|
24 (17) |
Leaf blade (4–)8–16 × 3–9 cm, base shallowly cordate to rounded; corolla broadly campanulate,
6–13 mm. |
|
(25) |
+ |
Leaf blade 1.7–11 × 1–4 cm (not G. dumicola), base usually cuneate or obtuse-rounded; corolla
urceolate or tubular, rarely campanulate, (2–)3–5(–6) mm. |
|
(27) |
|
|
|
|
25 (24) |
Corolla red; inflorescence shortly corymbose-racemose, 2–8 mm, glabrous. |
|
4 Gaultheria pseudonotabilis |
+ |
Corolla white; inflorescence racemose or corymbose, 10–60(–100) mm, densely tomentulose or glabrous. |
|
(26) |
|
|
|
|
26 (25) |
Corolla 1–1.3 cm; bracteoles basal; inflorescence densely tomentulose. |
|
5 Gaultheria codonantha |
+ |
Corolla 6–7 mm; bracteoles apical; inflorescence glabrous. |
|
21 Gaultheria leucocarpa |
|
|
|
|
27 (24) |
Leaf blade (4.5–)6–17 × 2.5–9 cm; inflorescence shortly racemose or corymbose, 3–10 mm. |
|
2 Gaultheria dumicola |
+ |
Leaf blade 1.7–11 × 1–4 cm; inflorescence racemose, at least 10 mm. |
|
(28) |
|
|
|
|
28 (27) |
Dwarf shrub, (5–)10–25 cm tall; leaf blade elliptic or broadly ovate, 1.7–6 × 1–4 cm, base obtuse to shallowly cordate; raceme 1–2.5 cm. |
|
12 Gaultheria praticola |
+ |
Shrub more than 30 cm tall; leaf blade elliptic to lanceolate, sometimes ovate, 3–11 × 1–4 cm, base attenuate to obtuse or rounded; raceme 1–5(–7) cm. |
|
(29) |
|
|
|
|
29 (28) |
Bracts orbicular-ovate or oblong, 4–7 mm; bracteoles longer than bracts or not. |
|
(30) |
+ |
Bracts ovate or triangular-ovate, 1–3(–4) mm; bracteoles smaller than bracts. |
|
(31) |
|
|
|
|
30 (29) |
Bracts 5–7 mm; bracteoles smaller than bracts, subapical or at middle of pedicel; leaf blade
abaxially brown, secondary veins (3 or)4(–7) pairs; twigs densely hirsute. |
|
10 Gaultheria hookeri |
+ |
Bracts ca. 4 mm; bracteoles 4–5.5 mm, apical; leaf blade abaxially grayish white or pale
brown, secondary veins 2 or 3 pairs; twigs densely hispid and puberulous. |
|
15 Gaultheria longibracteolata |
|
|
|
|
31 (29) |
Flowers 4-merous; leaf blade ca. 2 × as long as wide. |
|
14 Gaultheria tetramera |
+ |
Flowers 5-merous; leaf blade 2.5–4 × as long as wide. |
|
(32) |
|
|
|
|
32 (31) |
Ovary half-inferior; stamens 5. |
|
17 Gaultheria semi-infera |
+ |
Ovary superior; stamens 10. |
|
(33) |
|
|
|
|
33 (32) |
Leaf blade abaxially stramineous or grayish white, appressed setulose; pedicel 1–2 mm. |
|
13 Gaultheria straminea |
+ |
Leaf blade abaxially densely long setulose and puberulous; pedicel 3–6 mm. |
|
1 Gaultheria wardii |
|