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3. ROSACEAE Subfam. ROSOIDEAE

蔷薇亚科 qiang wei ya ke

Shrubs or herbs. Leaves stipulate; leaf blade compound, rarely simple, margin usually serrate. Carpels many, free; ovary superior, rarely inferior; ovules 1 or 2, pendulous or erect, anatropous. Fruit an achene (torus sometimes fleshy and berrylike), rarely a drupelet. x = 7, 8, 9.

About 35 genera and 1500 species: nearly cosmopolitan, especially in temperate and subarctic regions of N hemisphere; 22 genera (one endemic) and 457 species (240 endemic) in China.


1 Achenes enclosed in cupular or urn-shaped torus.   (2)
+ Achenes or drupelets borne on flat or convex torus.   (7)
       
2 (1) Carpels numerous; torus fleshy at maturity; shrubs; stems usually prickly.   41 Rosa
+ Carpels 1–4; torus dry and hard at maturity; herbs perennial or shrublets; stems not prickly.   (3)
       
3 (2) Flowers 3-merous; style basal; shrublets.   42 Potaninia
+ Flowers (4 or)5-merous; style terminal, rarely basal or sub-basal; herbs perennial.   (4)
       
4 (3) Petals present.   (5)
+ Petals absent.   (6)
       
5 (4) Hypanthium with hooklike spines abaxially; epicalyx segments absent; stamens 5–15.   43 Agrimonia
+ Hypanthium without hooklike spines abaxially; epicalyx segments present; stamens 35–40.   44 Spenceria
       
6 (4) Leaves pinnate; sepals imbricate, epicalyx segments absent; stamens 4–15; style terminal; flowers usually in spicate or capitate inflorescences, bisexual, rarely unisexual (when plants monoecious).   45 Sanguisorba
+ Leaves simple, mostly palmately lobed or parted; sepals valvate, epicalyx segments present; stamens 1–5; style basal or sub-basal; flowers usually in corymbose cymes, always bisexual.   46 Alchemilla
       
7 (1) Carpels 4–15, borne at base of flat or slightly concave torus; stipules not adnate to petiole.   (8)
+ Carpels several to numerous, borne on globose or conic torus; stipules adnate to petiole or not.   (10)
       
8 (7) Herbs perennial; leaves pinnate or pinnately or palmately lobed; flowers numerous in a terminal panicle or corymb, small.   27 Filipendula
+ Shrubs; leaves simple; flowers solitary, large.   (9)
       
9 (8) Leaves alternate; flowers 5-merous; petals yellow; epicalyx segments absent; carpels 5–8.   25 Kerria
+ Leaves opposite; flowers 4-merous; petals white; epicalyx segments present; carpels 4.   26 Rhodotypos
       
10 (7) Fruit drupelets or drupaceous achenes; stems prickly, rarely unarmed.   28 Rubus
+ Fruit achenes; stems unarmed.   (11)
       
11 (10) Style terminal or subterminal.   (12)
+ Style basal, lateral, or subterminal.   (17)
       
12 (11) Style deciduous.   (13)
+ Style persistent.   (14)
       
13 (12) Herbs low; basal leaves pinnate, with many leaflets; carpels many; stamens persistent.   33 Coluria
+ Herbs creeping; basal leaves 3–5-foliolate or 3–5-parted; carpels 2–6; stamens deciduous.   34 Waldsteinia
       
14 (12) Shrubs dwarf; leaves simple, entire or lobed; flowers (6–)8–10-merous.   29 Dryas
+ Herbs perennial; basal leaves pinnate, rarely simple; flowers 5-merous.   (15)
       
15 (14) Style jointed, distal part separating from joint at fruit maturity, proximal part persistent and apically hooked.   30 Geum
+ Style not jointed, wholly persistent at fruit maturity.   (16)
       
16 (15) Basal leaves pinnate, with many leaflets; plants hermaphroditic; petals yellow; style not or slightly elongated at fruiting.   31 Acomastylis
+ Basal leaves simple; plants polygamo-monoecious or polygamo-dioecious; petals white; style distinctly elongated at fruiting.   32 Taihangia
       
17 (11) Torus inflated and fleshy when ripe.   (18)
+ Torus dry when ripe.   (19)
       
18 (17) Petals white; epicalyx segments smaller than sepals.   39 Fragaria
+ Petals yellow; epicalyx segments larger than sepals.   40 Duchesnea
       
19 (17) Stamens and carpels many.   (20)
+ Stamens 4 or 5; carpels 4–20.   (21)
       
20 (19) Petals yellow or white, longer than sepals.   35 Potentilla
+ Petals purple or white, shorter than or subequaling sepals.   36 Comarum
       
21 (19) Stamens alternate with petals; basal leaves palmately or pinnately compound; epicalyx segments present.   37 Sibbaldia
+ Stamens opposite to petals; leaves 3-parted or 2 or 3 × compound; epicalyx segments absent.   38 Chamaerhodos

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