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25. Tribe GALEGEAE
山羊豆族 shan yang dou zu
Authors: Langran Xu, Xiangyun Zhu, Bojian Bao, Mingli Zhang, Hang Sun, Dietrich Podlech, Stanley L. Welsh, Hiroyoshi Ohashi, Kai Larsen & Anthony R. Brach
Herbs or shrubs, with simple or T-shaped hairs; glands or glandular punctae sometimes present. Leaves epulvinate or pulvinus reduced, imparipinnate or paripinnate, with many opposite to irregularly arranged or rarely conjugate leaflets, rarely 1-3-foliolate; stipules free or adnate to petiole, estipellate. Flowers in axillary racemes, spikes, or rarely solitary. Calyx campanulate to tubular; standard clawed or narrowed to base; wings auriculate; keel blunt to apiculate. Stamens diadelphous, rarely monadelphous; anthers usually uniform, but slightly dimorphic and with confluent thecae in Glycyrrhiza. Ovary few to many ovuled (sometimes 1-seeded); style slender, bearded or not, with a terminal or lateral stigma. Legumes compressed, angled or inflated, sometimes with sutured margins intruded or longitudinally septate, occasionally torulose, dehiscent or indehiscent. Seeds oblong-reniform, estrophiolate.
About 24 genera and 2900-3200 species: principally in Asia, Europe, and North America, but extending thinly in mountainous and/or drier places to S Africa, Australia, and temperate South America; 11 genera and 586 species (324 endemic, two introduced) in China.
Galega officinalis Linnaeus (Sp. Pl. 2: 714. 1753), probably native to SW Asia (Caucasus), is cultivated in China.
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| 1 |
Style bearded, sometimes just a tuft of hairs below stigma on one side; wings and keel never interlocking (subtribe Coluteinae) |
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(2) |
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Style glabrous, but stigma sometimes penicillate (sometimes inner side of style upper part hairy in Astragalus); wings and keel mostly interlocking with bosses on keel and matching sockets on wings (indistinct only in Gueldenstaedtia and Tibetia) |
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(5) |
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| 2 (1) |
Leaves reduced to scales; flowers solitary; legumes compressed. |
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145 Eremosparton |
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Leaves imparipinnate, 7-25-foliolate; flowers in racemes; legumes inflated |
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(3) |
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| 3 (2) |
Flowers yellow; standard with 2 callosities above claw; shrubs. |
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143 Colutea |
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Flowers red to violet, rarely yellow; standard without callosities above claw; shrubs or herbs |
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(4) |
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| 4 (3) |
Plants with medifixed hairs; legumes inflated subglobose. |
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144 Sphaerophysa |
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Plants with basifixed hairs; legumes narrowly ellipsoid to ovoid. |
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138 Phyllolobium |
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| 5 (1) |
Anther thecae confluent toward apex; leaves pulvinate, spirally arranged; some hairs glandular-peltate or -capitate (subtribe Glycyrrhizinae). |
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148 Glycyrrhiza |
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Anther thecae separate; leaf pulvinus reduced or absent; plants without peltate glandular hairs |
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(6) |
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| 6 (6) |
Legume nerves oblique, closely parallel, raised at maturity, confluent toward placental margin; vexillary filament connate with others; keel petals not auriculate; leaflet nerves extending to margin (subtribe Galeginae). |
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Galega (see note above)
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Legume nerves transverse, confluent over middle part of valves, fine or inconspicuous; vexillary filament generally free; keel petals generally auriculate; leaflet nerves usually looped within margin or inconspicuous (subtribe Astragalinae) |
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(7) |
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| 7 (6) |
Calyx base usually oblique, upper side ± slightly gibbous; wings pinnately nerved; legume valves generally twisting on dehiscing |
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(8) |
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Calyx base symmetric or suboblique; wings usually palmately nerved; legume valves not twisting |
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(9) |
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| 8 (7) |
Stem obvious; stipules membranous, separated from petiole; calyx campanulate. |
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142 Chesniella |
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Stem abbreviated, plants appearing acaulescent; stipules herbaceous, adnate to petiole; calyx tubular. |
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141 Chesneya |
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| 9 (7) |
Keel petals half as long as wings; style shorter than or as long as ovary |
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(10) |
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Keel petals subequal to or slightly shorter than wings; style longer than ovary |
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(11) |
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| 10 (9) |
Style incurved; upper 2 calyx lobes free; seeds scrobiculate; stipules free and adnate to base of petiole. |
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146 Gueldenstaedtia |
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Style curved in right angle; upper 2 calyx lobes connate; seeds smooth, marbled-spotted; stipules connate below apex and opposite to leaves. |
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147 Tibetia |
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| 11 (9) |
Keel petals obtuse; legume 1-locular or with a septum intruding from abaxial suture; leaflets ± symmetric. |
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139 Astragalus |
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Keel petals apiculate; legume 1-locular or with a septum intruding from adaxial suture; leaflets basally ± oblique or, if narrow, falcately incurved. |
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140 Oxytropis |
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List of lower taxa
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