| 
              
                
                  
                    
                      
                      
                      
	
                        
8. Polygonum Linnaeus, Sp. Pl.  359.  1753; Gen. Pl. ed. 2. 116. 1754 (p.p.); Hook. f., Fl. Brit. Ind. 5: 23 1886 (p.p.); R.N.Parker, For. Fl. Punj. Haz. & Delhi 423. 1921 p.p. (reprint. ed.); D.A.Webb & Chater in Tutin et al., Fl. Europ. 1: 76. 1964 (p.p.); Rech. f. & Schiman-Czeika in Rech. f., Fl. Iran. 56: 46. 1968 (p.p.); Bhopal & Chaudhri in Pak. Syst. 1: 66. 1977 (p.p.); Hara in Hara et al., Enum. Fl. Pl. Nep. 3: 178. 1982; Grierson & D.G.Long, ; Fl. Bhutan 1(1): 170. 1983; Munshi & Javeid, Syst. Stud. Polyg. Kashm. Him. 52. 1986; Chaudhary, Fl. Kingd. Saudi Arab. 1: 305. 1999; Boulos, Fl. Egypt 1: 27. 1999.  
 
 
 
 
Annual or perennial, prostrate, subprostrate to erect, branched or simple herbs. Stem 1-many, branched from base or above. Leaves small, linear-lanceolate, ovate or elliptic, sessile-petiolate. Ochreae small, silvery or whitish hyaline membranous, partite or ciliate-lacerate. Inflorescence axillary, few flowered or solitary, or in small clusters. Perianth mostly petaloid, tepals 4-6, elliptic-ovate. Stamens 4-8, filaments short, usually broadened at the base without any interstaminal nectaries. Ovary biconvex or trigonous, small, with 2-3 styles and capitate stigmas. Nuts biconvex or trigonous, dark brown-black, shining. 
A genus (s. str.) of c. 60 species, distributed in both the hemispheres. Represented by 20 species in Pakistan. 
 
 
 
 
 
 
 
 
                        
                         
		
                          
                          
			
				| 
                                 
                                   
                               | 
				
                                 
                                   
                               | 
			 
			
				| 
                                 
                                   
                                
                                  
  | 1 | 
  Plants annual. | 
    | 
  (10) | 
 
  | + | 
  Plants perennial. | 
    | 
  (2) | 
 
  |   | 
    | 
    | 
    | 
 
  | 2 (1) | 
  Ochreae membranous or somewhat hyaline, conspicuously nerved throughout or more than half of their length. | 
    | 
  (3) | 
 
  | + | 
  Ochreae hyaline, not conspicuously nerved or nerved at the base only. | 
    | 
  (6) | 
 
  |   | 
    | 
    | 
    | 
 
  | 3 (2) | 
  Nerves of the ochreae thickened and exserted like 2 aristae. | 
    | 
  1 P. biaristatum | 
 
  | + | 
  Nerves of the ochreae though strong but not thickened and aristate, rarely exserted. | 
    | 
  (4) | 
 
  |   | 
    | 
    | 
    | 
 
  | 4 (3) | 
  Ochreae longer or almost equalling the internodes, covering the internodes, with 12 strong nerves. Nuts 2.5-3.5 mm long. | 
    | 
  4 P. maritimum | 
 
  | + | 
  Ochreae shorter than the internodes, not covering the internodes, with 2 nerves only. Nuts 1.5-2 mm long. | 
    | 
  (5) | 
 
  |   | 
    | 
    | 
    | 
 
  | 5 (4) | 
  Ochreae shorter than the internodes, not covering the internodes, with 2 nerves only. Nuts 1.5-2 mm long. | 
    | 
  2 P. recumbens | 
 
  | + | 
  Plants prostrate. Leaf blades linear, gland-dotted. Nuts dull and striate. | 
    | 
  3 P. roylei | 
 
  |   | 
    | 
    | 
    | 
 
  | 6 (2) | 
  Leaves sessile, linear lanceolate. | 
    | 
  (7) | 
 
  | + | 
  Leaves petiolate, elliptic, oblong to oblong-spathulate, ovate. | 
    | 
  5 P. cognatum | 
 
  |   | 
    | 
    | 
    | 
 
  | 7 (6) | 
  Plants erect. Rhizome distinctly many branched. Nuts ± symmetrical, regular, often trigonous, acute to pointed at the tip (leaves Thymus-like). | 
    | 
  6 P. thymifolium | 
 
  | + | 
  Plants prostrate to subprostrate. Rhizome unbranched. Nuts asymmetrical, distinctly curved, not pointed at the tip (leaves not Thymus-like). | 
    | 
  (8) | 
 
  |   | 
    | 
    | 
    | 
 
  | 8 (7) | 
  Flowering branches numerous, filiform, ± ascending, with few leaves. Leaves 0.5 mm broad. Ochreae ± 3 mm long. | 
    | 
  7 P. sarobiense | 
 
  | + | 
  Flowering branches few, thick, with many leaves. Leaves 1-2 (-3) mm broad. Ochreae 5-15 mm long. | 
    | 
  (9) | 
 
  |   | 
    | 
    | 
    | 
 
  | 9 (8) | 
  Leaves up to 10 (-12) mm long, ochrea always equalling or exceeding the leaves especially on young branches. Plants usually glabrous. | 
    | 
  8 P. paronychioides | 
 
  | + | 
  Leaves 10-25 mm long. Ochrea always shorter than the leaves. Plants papillose-puberulous. | 
    | 
  9 P. afghanicum | 
 
  |   | 
    | 
    | 
    | 
 
  | 10 (1) | 
  Plants very small, stem dichotomously branched, up to 5 cm long. Perianth hyaline. Ochrea almost covering the flowers. | 
    | 
  20 P. molliaeforme | 
 
  | + | 
  Plants large. Stem >5 cm long not dichotomously branched. Perianth petaloid, white, green, roseate or red. Flowers not covered by ochreae. | 
    | 
  (11) | 
 
  |   | 
    | 
    | 
    | 
 
  | 11 (10) | 
  Flower bearing branches either without leaves or bracteal leaves much smaller thus the floral branches conspicuously differentiated from the leafy vegetative branches. | 
    | 
  (12) | 
 
  | + | 
  Flowers bearing branches with normal leaves and the flowers ± disposed over the entire plant almost from the base. | 
    | 
  (13) | 
 
  |   | 
    | 
    | 
    | 
 
  | 12 (11) | 
  Inflorescence compact. Floral branches almost naked. Tepals parted almost half the length. Nuts 1.5-2.0 mm long. | 
    | 
  10 P. argyrocoleum | 
 
  | + | 
  Inflorescence lax. Floral branches not completely naked. Tepals parted almost up to the base. Nuts 2-3 mm long. | 
    | 
  11 P. patulum | 
 
  |   | 
    | 
    | 
    | 
 
  | 13 (11) | 
  Internodes ± covered by the ochreae and closely imbricated leaves. | 
    | 
  (14) | 
 
  | + | 
  Internodes not covered by the ochreae and closely imbricated leaves. | 
    | 
  (16) | 
 
  |   | 
    | 
    | 
    | 
 
  | 14 (13) | 
  Plants mostly glabrous. Perianth subcampanulate, tube shorter than the lobes usually not fused with nuts. Nuts not shining. | 
    | 
  (15) | 
 
  | + | 
  Plants mostly puberulous. Perianth tubular, tube longer than the lobes, usually fused with nuts. Nuts shining. | 
    | 
  19 P. rottboellioides | 
 
  |   | 
    | 
    | 
    | 
 
  | 15 (14) | 
  Perianth scarcely 1 mm long, exceeded by the mature nuts. Leaves membranous, oval. | 
    | 
  17 P. cashmeriense | 
 
  | + | 
  Perianth c. 2 mm long, equalling to mature achenes. Leaves coriaceous, linear to ovate. | 
    | 
  18 P. polycnemoides | 
 
  |   | 
    | 
    | 
    | 
 
  | 16 (13) | 
  Plants heterophyllous. Cauline leaves larger, ramal leaves smaller. | 
    | 
  (17) | 
 
  | + | 
  Plants not heterophyllous. Cauline and ramal leaves ± equal size. | 
    | 
  (18) | 
 
  |   | 
    | 
    | 
    | 
 
  | 17 (16) | 
  Nuts smooth. Stem and branches not striate. | 
    | 
  14 P. olivascens | 
 
  | + | 
  Nuts rugolose. Stem and branches mostly striate. | 
    | 
  12 P. aviculare | 
 
  |   | 
    | 
    | 
    | 
 
  | 18 (16) | 
  Plants erect, ochrea distinctly nerved from the base to the apex. | 
    | 
  11 P. patulum | 
 
  | + | 
  Plants prostrate to procumbent, ochrea either not nerved or faintly nerved. | 
    | 
  (19) | 
 
  |   | 
    | 
    | 
    | 
 
  | 19 (18) | 
  Ochrea deeply bifid. Nuts 2-2.5 mm long. | 
    | 
  13 P. arenastarum | 
 
  | + | 
  Ochrea undivided, mostly lacerate or toothed only at the apex. Nuts 1-1.25 (1.5) mm long. | 
    | 
  (20) | 
 
  |   | 
    | 
    | 
    | 
 
  | 20 (19) | 
  Flowers distinctly pedicellate, exserted from the ochreae.All tepals almost equal in size. | 
    | 
  15 P. effusum | 
 
  | + | 
  Flowers sessile to subpedicellate, mostly sunken among the ochreae, rarely exserted. Outer tepals slightly longer. | 
    | 
  16 P. plebejum | 
 
 
 
                               | 
			 
			
				| 
                                 | 
			 
		 
		
                        
	  
                        
                          List of lower taxa 
 
                        
                          
                        
	
                        
                      
 
                      
	 
                      
                         
		
			| Related Links (opens in a new window) | 
		 
		
			| 
                               Treatments in Other Floras @ www.efloras.org
Other Databases
 
 
                             | 
		 
	 
	
                        
 
                     | 
                   
                  
                    | 
                      
                      
                        
                      
                         
                     | 
                   
                 
               | 
               | 
             
           
         |