1 |
Spikelets strongly laterally compressed, lemmas 7–13-veined, keeled (B. sect. Ceratochloa (P. Beauvois) Grisebach) |
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(2) |
+ |
Spikelets slightly laterally compressed to ± cylindrical, lemmas 5–9-veined, rounded on back |
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(4) |
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2 (1) |
Perennial; leaf sheaths retrorsely pubescent; lemmas often hairy, awn 5–7 mm. |
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53 B. marginatus |
+ |
Annual; leaf sheaths glabrous; lemmas mostly glabrous, mucronate or awned |
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(3) |
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3 (2) |
Lemmas awnless or with mucro, ca. 1 mm; palea 1/2 lemma length. |
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54 B. catharticus |
+ |
Lemmas with ca. 10 mm awn; palea subequal to lemma. |
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55 B. carinatus |
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4 (1) |
Perennials, tufted or rhizomatous |
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(5) |
+ |
Annuals |
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(37) |
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5 (4) |
Glumes subequal to lower florets; awn stout, recurved, mostly as long as lemma; panicle narrow, branches simple (B. sect. Sinobromus Keng ex P. C. Keng) |
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(6) |
+ |
Glumes conspicuously shorter than lower florets; awn slender, straight, mostly shorter than lemma; panicle contracted to effuse (B. sect. Bromopsis Dumortier) |
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(10) |
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6 (5) |
Lower glume 5–8 mm, upper glume 7–12 mm; lemmas 8–10 mm. |
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24 B. himalaicus |
+ |
Lower glume 8–15 mm, upper glume 11–17 mm; lemmas 9–16 mm |
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(7) |
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7 (6) |
Glumes glabrous |
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(8) |
+ |
Glumes pubescent |
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(9) |
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8 (7) |
Culms 50–70 cm tall, nodes 3; lemmas glabrous. |
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26 B. epilis |
+ |
Culms up to 100 m tall, nodes 7–8; lemmas marginally and distally pubescent. |
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27 B. mairei |
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9 (7) |
Panicle branches each with 1–5 spikelets; spikelets 12–20 mm; awn arising near lemma apex. |
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28 B. sinensis |
+ |
Panicle branches each with 1 apical spikelet; spikelets 18–25 mm; awn arising between apical lobes. |
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25 B. porphyranthos |
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10 (5) |
Plants usually more than 1 m tall; lemma with long or short awn |
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(11) |
+ |
Plants less than 1 m tall (if more than 1 m, then lemma awn shorter than 3 mm, sometimes absent) |
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(24) |
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11 (10) |
Plants tufted, without rhizomes or stolons; culms with 6–9 nodes |
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(12) |
+ |
Plants rhizomatous; culms often with fewer than 6 nodes |
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(16) |
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12 (11) |
Lemma 3–5-veined; rachilla clearly visible from side in mature spikelet |
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(13) |
+ |
Lemma often 7-veined; rachilla not visible from side of intact mature spikelet |
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(14) |
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13 (12) |
Rachilla internodes 3–4 mm; lemma 5-veined, ca. 12 mm. |
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1 B. magnus |
+ |
Rachilla internodes 1–2 mm; lemma 3-veined, ca. 10 mm. |
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2 B. plurinodis |
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14 (12) |
Panicles contracted; lemmas 10–12 mm. |
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3 B. erectus |
+ |
Panicles spreading; lemmas 12–15 mm |
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(15) |
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15 (14) |
Panicle branches 3–5 cm, with 1–3 spikelets. |
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4 B. pseudoramosus |
+ |
Panicle branches up to 20 cm, with 2–9 spikelets. |
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5 B. ramosus |
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16 (11) |
Plants with long creeping rhizomes |
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(17) |
+ |
Plant with short rhizomes |
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(20) |
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17 (16) |
Panicles spreading. |
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7 B. pumpellianus |
+ |
Panicles narrow |
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(18) |
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18 (17) |
Panicles racemelike; lemmas glabrous. |
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6 B. stenostachyus |
+ |
Panicles narrowly contracted; lemmas pubescent on veins |
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(19) |
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19 (18) |
Lemma pubescent along keel and marginal veins to middle or slightly above; awn 1–5 mm. |
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8 B. sibiricus |
+ |
Lemma pubescent on marginal veins almost to apex; awn up to 2 mm. |
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14 B. korotkiji |
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20 (16) |
Panicle branches (2–)6–9 per node |
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(21) |
+ |
Panicle branches 2–5 per node |
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(22) |
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21 (20) |
Awns 1–1.5 mm; rachilla not visible from side, internodes very short. |
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13 B. inermis |
+ |
Awns 5–10 mm; rachilla clearly visible from side, internodes 3–4 mm. |
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12 B. remotiflorus |
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22 (20) |
Leaf blades 3–5 mm wide. |
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9 B. staintonii |
+ |
Leaf blades 5–12 mm wide |
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(23) |
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23 (22) |
Lemmas 11–14 mm; anthers 2.5–3 mm. |
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10 B. benekenii |
+ |
Lemmas 10–12 mm; anthers 1–1.2 mm. |
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11 B. ciliatus |
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24 (10) |
Awns absent or less than 2(–4) mm; plants with long rhizomes |
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(25) |
+ |
Awns more than 2 mm; plants tufted or rhizomatous |
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(26) |
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25 (24) |
Lemmas glabrous, scabrid near base; anthers 3–4 mm. |
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13 B. inermis |
+ |
Lemmas pilose, at least on keel and marginal veins; anthers 5–6 mm. |
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14 B. korotkiji |
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26 (24) |
Plants with long downwardly extending rhizomes. |
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15 B. formosanus |
+ |
Plants tufted or with short rhizomes |
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(27) |
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27 (26) |
Plants loosely tufted, or with short inconspicuous rhizomes |
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(28) |
+ |
Plants densely tufted |
|
(32) |
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28 (27) |
Plants ca. 20 cm tall; panicle contracted, lower branches crowded. |
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20 B. variegatus |
+ |
Plants 20–60 cm tall; panicle open, lower branches not crowded, spreading |
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(29) |
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29 (28) |
Plants ca. 60 cm tall, with short rhizomes. |
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21 B. confinis |
+ |
Plants ca. 20–50 cm tall, tufted |
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(30) |
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30 (29) |
Awns ca. 3 mm. |
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17 B. paulsenii |
+ |
Awns 8–12 mm |
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(31) |
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31 (30) |
Panicle branches each with 2–4 spikelets; spikelets 2–2.5 cm. |
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22 B. morrisonensis |
+ |
Panicle branches each with 1 or 2 spikelets; spikelets 3–4 cm. |
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23 B. nepalensis |
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32 (27) |
Plants 40–90 cm tall |
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(33) |
+ |
Plants up to 40(–50) cm tall |
|
(34) |
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33 (32) |
Lower glume 8–10 mm, upper glume 10–12 mm. |
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17 B. paulsenii |
+ |
Lower glume 5–6 mm, upper glume 7–8 mm. |
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19 B. tyttholepis |
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34 (32) |
Awns 6–8 mm. |
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16 B. riparius |
+ |
Awns 3–5 mm |
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(35) |
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35 (34) |
Leaf blades ca. 5 mm wide; panicles 10–12 cm. |
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17 B. paulsenii |
+ |
Leaf blades 1.5–2 mm wide; panicles 5–9 cm |
|
(36) |
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36 (35) |
Panicle branches scabrid, shorter than spikelet; lemmas glabrous, often scabrid. |
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17 B. paulsenii |
+ |
Panicle branches slender, flexuous, often longer than spikelets; lemmas pubescent. |
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18 B. cappadocicus |
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37 (4) |
Spikelets broadened upward, wedge-shaped; glumes narrow, lower glume 1(–3)-veined, upper glume 3(–5)-veined |
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(38) |
+ |
Spikelets narrowed upward, oblong-lanceolate; glumes broad, lower glume 3–5-veined, upper glume 5–9-veined (B. sect. Bromus) |
|
(44) |
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38 (37) |
Awn 4–6 times length of lemma; lemma apex minutely 2-toothed (B. sect. Nevskiella (Kreczetowicz & Vvedensky) Tournay). |
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35 B. gracillimus |
+ |
Awn 1–3 times length of lemma; lemma apex clearly 2-toothed, teeth 3–7 mm (B. sect. Genea Dumortier) |
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(39) |
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39 (38) |
Panicle lax, branches spreading or drooping |
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(40) |
+ |
Panicle contracted, branches erect or ascending, never drooping |
|
(41) |
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40 (39) |
Panicle much branched, branches longer or shorter than spikelets, pubescent; spikelets 1–8 per branch, secund. |
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29 B. tectorum |
+ |
Panicle simple, branches up to 10 cm, with 1 or no branchlets; spikelets sparse, 1 or 2(or 3) per branch. |
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30 B. sterilis |
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41 (39) |
Lower glume 15–20 mm; lemmas 20–30 mm, awn 30–40 mm. |
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31 B. rigidus |
+ |
Lower glume 6–10 mm; lemmas 10–15 mm, awn 10–20 mm |
|
(42) |
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42 (41) |
Panicle lax; branches 2–3 cm, scabrid; culms glabrous below inflorescence; stamens 2. |
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32 B. madritensis |
+ |
Panicle contracted; branches shorter than spikelets, ca. 1 cm, culms pubescent below inflorescence; stamens usually 3 |
|
(43) |
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43 (42) |
Plants 5–15 cm tall; panicle ca. 5 cm, branches each with 1 or 2 spikelets; lemmas 13–15 × 1–1.5 mm. |
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33 B. fasciculatus |
+ |
Plants 30–40 cm tall; panicle ca. 10 cm, branches each with 4 or 5 spikelets; lemmas 10–13 × ca. 2 mm. |
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34 B. rubens |
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44 (37) |
Lemmas 3-awned. |
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52 B. danthoniae |
+ |
Lemmas 1-awned |
|
(45) |
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45 (44) |
Spikelets (2–)3–4 cm; lemmas 14–18 mm |
|
(46) |
+ |
Spikelets (1–)1.5–2.5(–3) cm; lemmas 5–11 mm |
|
(49) |
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46 (45) |
Spikelets oblong-lanceolate, ca. 6 mm wide; lemma apex deeply toothed, teeth (1.5–)3–4 mm; panicle open, branches several times longer than spikelets, nodding |
|
(47) |
+ |
Spikelets broadly elliptic, ca. 10 mm wide; lemma apex shallowly toothed, teeth 1–2 mm; panicle slightly contracted, branches not longer than spikelets, obliquely ascending |
|
(48) |
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47 (46) |
Lower glume ca. 10 mm; lemma 15–18 mm, awn 20–25 mm, lower part slightly flattened, twisted, conspicuously recurved |
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48 B. oxyodon |
+ |
Lower glume 5–8 mm; lemma 10–15 mm, awn 8–17 mm, terete, straight, usually erect. |
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49 B. pectinatus |
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48 (46) |
Lemma scabrid; awn 5–10 mm, straight; anthers 2–3 mm; panicle branches on 1 side of axis. |
|
50 B. grossus |
+ |
Lemma smooth; awn 10–20 mm, conspicuously recurved; panicle branches on all sides on axis. |
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51 B. lanceolatus |
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49 (45) |
Lemma margins inrolled, not covering adjacent florets; rachilla internodes exposed in mature spikelets. |
|
36 B. secalinus |
+ |
Lemma margins overlapping adjacent florets; rachilla internodes enclosed in mature spikelets |
|
(50) |
|
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50 (49) |
Lemmas 5–7 mm; awn shorter than lemma body, straight; spikelets 1–1.5(–2.5) cm |
|
(51) |
+ |
Lemmas 8–12 mm, awn as long as or longer than lemma body, straight or recurved; spikelets (1.5–)2–3 cm |
|
(53) |
|
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51 (50) |
Lemmas 6–11 mm; anthers 1.5–3 mm; caryopsis enclosed by lemma, not exposed. |
|
41 B. racemosus |
+ |
Lemmas 5–6 mm; anthers 0.5–1.5 mm; caryopsis longer than lemma, exposed |
|
(52) |
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52 (51) |
Culms 30–50 cm tall; leaf blades ca. 10 cm; panicle 5–10 cm; lemmas 5.5–6.5 mm. |
|
37 B. lepidus |
+ |
Culms ca. 10 cm tall; leaf blades 3–4 cm; panicle ca. 3 cm; lemmas 4–5 mm. |
|
38 B. brachystachys |
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53 (50) |
Lemmas broadly ovate-elliptic, 3–4 mm wide in side view, margins broadly membranous with a conspicuous obtuse angle about halfway between base and apex; panicle subracemose, branches each with 1 or 2 apical spikelets. |
|
39 B. squarrosus |
+ |
Lemmas elliptic, ca. 2 mm wide in side view, margins without any conspicuous angle; panicle laxly spreading or contracted |
|
(54) |
|
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54 (53) |
Panicle lax, open, 15–25 cm, branches and pedicels longer than spikelets |
|
(55) |
+ |
Panicle contracted, 3–10 cm, branches and pedicels shorter than spikelets |
|
(58) |
|
|
|
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55 (54) |
Anthers ca. 4 mm; palea subequal to lemma; leaf sheaths glabrous. |
|
40 B. arvensis |
+ |
Anthers 0.5–3 mm; palea conspicuously shorter than lemma; leaf sheaths pubescent |
|
(56) |
|
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56 (55) |
Awns straight; all awns within a spikelet subequal. |
|
41 B. racemosus |
+ |
Awns curved outward, lower awns within a spikelet shorter than upper awns |
|
(57) |
|
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57 (56) |
Lemma 10–15 mm, apex deeply 2-lobed, acuminate; awn arising 3–4 mm below apex, terete, spreading. |
|
49 B. pectinatus |
+ |
Lemma 8–10 mm, apex with short, obtuse teeth; awn arising 1–2 mm below apex, base slightly flattened, recurved outward when mature. |
|
42 B. japonicus |
|
|
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|
58 (54) |
Lemmas glabrous; awn as long as or slightly longer than lemma body |
|
(59) |
+ |
Lemmas pubescent; awn slightly shorter than lemma body |
|
(61) |
|
|
|
|
59 (58) |
Panicle ca. 10 cm; lower glume ca. 7 mm, awn straight, arising ca. 1 mm below lemma apex; anthers ca. 1 mm. |
|
43 B. sewerzowii |
+ |
Panicle 3–7 cm; lower glume 3–4 mm; awn curved downward, arising 1.5–3 mm below lemma apex; anthers 0.3–0.6 mm |
|
(60) |
|
|
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60 (59) |
Panicle with few spikelets, 1-sided; lemmas 8–10 mm, apical teeth 0.5–1 mm. |
|
44 B. tytthanthus |
+ |
Panicle dense, with many clustered spikelets; lemmas 7–8 mm, apical teeth 1.3–2.5 mm. |
|
45 B. scoparius |
|
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|
61 (58) |
Plant 40–80 cm tall; culms pubescent on nodes and below panicle, panicle branches and pedicels pubescent; awns slender, straight. |
|
46 B. hordeaceus |
+ |
Plant 20–40(–60) cm tall; culms, panicle branches, and pedicels glabrous; awns conspicuously recurved, base slightly flattened, twisted |
|
(62) |
|
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62 (61) |
Spikelets 10–15 mm; lower glume 2/3 as long as upper glume; lemma apex with 2–3 mm teeth; anthers ca. 1 mm. |
|
47 B. intermedius |
+ |
Spikelets 15–50 mm; glumes subequal; lemma apex with 1–1.5 mm teeth; anthers 1–1.5 mm. |
|
39 B. squarrosus |