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1. Subtribe TUSSILAGININAE  
款冬亚族 kuan dong ya zu 
 
Authors: Shangwu Liu, Yilin Chen, Irina D. Illarionova, Bertil Nordenstam, Hiroshige Koyama & Charles Jeffrey
 
 
 
 
Antheropodia cylindric or obconic, without enlarged marginal basal cells; endothecial tissue cell wall thickenings polarized, cells short; stigmatic areas usually confluent or contiguous; chromosome number usually 30, sometimes 29, 28, 27, 26, or 25, rarely 20 or 19. 
About 60 genera and 900 species: worldwide; 11 genera (three endemic) and 269 species (193 endemic) in China. 
 
 
 
 
 
 
 
 
                        
                         
		
                          
                          
			
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  | 1 | 
  Phyllaries in 2 series, all alike, herbaceous. | 
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  92 Doronicum | 
 
  | + | 
  Phyllaries in 1 series, herbaceous, cartilaginous, or leathery, if an outer series (calyculus) also present then distinct in size and form from inner series | 
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  (2) | 
 
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  | 2 (1) | 
  Inner florets functionally male; florets precocious | 
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  (3) | 
 
  | + | 
  Inner florets bisexual, florets not precocious | 
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  (4) | 
 
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  | 3 (2) | 
  Plants monoecious; peduncles 1-headed. | 
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  101 Tussilago | 
 
  | + | 
  Plants subdioecious; capitula with polygamous florets; peduncles several headed. | 
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  102 Petasites | 
 
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  | 4 (2) | 
  Leaves basally sheathed; achenes not beaked | 
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  (5) | 
 
  | + | 
  Leaves basally not sheathed; achenes beaked or not | 
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  (7) | 
 
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  | 5 (4) | 
  Leaf margins involute; achenes hairy. | 
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  93 Farfugium | 
 
  | + | 
  Leaf margins revolute; achenes glabrous | 
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  (6) | 
 
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  | 6 (5) | 
  Involucres cylindric or obconic. | 
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  94 Ligularia | 
 
  | + | 
  Involucres broadly campanulate or hemispheric. | 
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  95 Cremanthodium | 
 
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  | 7 (4) | 
  Capitula radiate; rhizomes dilated, tuberiform. | 
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  96 Sinacalia | 
 
  | + | 
  Capitula discoid, homogamous, florets bisexual; rhizomes not tuberiform | 
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  (8) | 
 
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  | 8 (7) | 
  Style branches with papillae divided at apex into 2 brushlike lobes. | 
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  97 Dicercoclados | 
 
  | + | 
  Style branches with papillae undivided at apex | 
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  (9) | 
 
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  | 9 (8) | 
  Cotyledons 1; radical leaves umbraculiform, pendulous when young. | 
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  100 Syneilesis | 
 
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  Cotyledons 2; basal leaves not umbraculiform, not pendulous when young | 
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  (10) | 
 
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  | 10 (9) | 
  Anthers caudate or sagittate; radical leaves withered by anthesis. | 
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  98 Parasenecio | 
 
  | + | 
  Anthers obtuse, not caudate; radical leaves persistent at anthesis. | 
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  99 Ligulariopsis | 
 
 
 
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                          List of lower taxa 
 
                        
                          
                        
                        
                      
 
                      
	 
                      
                         
		
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