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Pakistan | Family List | Brassicaceae

Sisymbrieae

Lateral nectar glands semiannular or annular, open towards the inner side, rarely in pairs or short; middle glands often joining the laterals, torose and ± thickened in the middle, rarely short, conical and not joining the laterals. Hairs simple or branched, sometimes absent, often rigid and ciliate, rarely glandular. Sepals erect or suberect, Stamen filaments not appendaged but sometimes broad at base; anthers usually oblong, obtuse. Stigma depressed capitate, often sub-bilobed. Fruits usually long (siliquae) rarely short (siliculae) ; valves often somewhat convex and rigid; style mostly short, rarely absent; stigma mostly broad, obscurely to sub-bilobed; septum usually complete, rarely perforated or absent,with or without a vein, rarely 2-4-veined; seeds usually oblong or suborbicular, rarely winged; mucilaginous or not when wet; radicle mostly incumbent.

Represented by 14 genera and c. 40 species in our area.

1 Leaves bipinnate   (2)
+ Leaves simple to lyrate-pinnatifed   (5)
       
2 (1) Perennial, caespitose herbs; fruits siliculae   Smelowskia
+ Annual or biennial herbs; fruits siliquae   (3)
       
3 (2) Pedicels very short, inconspicuous, thick; siliquae subangular, conical-tapering above   Robeschia
+ Pedicels conspicuous, often long, thin; fruits uniformly broad   (4)
       
4 (3) Fruits more than 12 mm long, linear, subcylin- drical, often upcurved   Descurainia
+ Fruits less than 12 mm long , narrowly ellipsoid often slightly narrowed below, usually straight   Sophiopsis
       
5 (1) Basal leaves ovate-cordate; seeds longitudinally striated, large   Alliaria
+ Basal leaves not so cordate; seeds not longitudinally striated, small   (6)
       
6 (5) Fruit short, siliculae (rarely dub-siliquae type)   (7)
+ Fruits often long, siliquae (rarely sub-siliculae type)   (9)
       
7 (6) Cauline leaves amplexicaul, often ± sagittate   Camelina
+ Cauline leaves linear to oblabceolate, not amplexicaul   (8)
       
8 (7) Cauline leaves several; racemes lax, mostly bracteate   Aphragmus
+ Cauline leaves 0-2 only; racemes often compact, ebracteate   Braya
       
9 (6) Radical leaves much larger than the cauline leaves, about as long as the aerial stem; seeds 2-seriate   Arcyosperma
+ Aerial stem usually much longer than the radical Leaves, seeds 1-seriate   (10)
       
10 (9) Plants tender, lax with filiform stem; leaves 3-lobed to entire; fruits short, 1-locular, 3-6-seeded   Lignariella
+ Plants erect to ascending, often robust; leaves not so lobed; fruits often long, 2-locular, many seeded   (11)
       
11 (10) Flowers yellow (in our species); lateral nectar glands hexagonal-quadrate; valves ± 3- veined   Sisymbrium
+ Flowers white, pinkish or violet (yellow in Arabidopsis pumila only) lateral nectar glands not angular; valves 1-veined   (12)
       
12 (11) Middle nectar glands absent   Torularia
+ Middle nectar glands present, ± torose, often joining the laterals   (13)
       
13 (12) Seeds not mucilaginous when wet   Microsisymbrium
+ Seeds usually mucilaginous when wet   Arabidopsis

  • List of lower taxa


     

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