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Pakistan | Family List | Papilionaceae | Astragalus

GROUP-B- (Astragalus)

Synopsis Of Sections And Species

Sect. Oxyglottis- Astragalus tribuloides, A. filicaulis, A. eremophilus, A. schimperi, A. oxyglottis

Sect. Poliothrix- Astragalus leucocephalus

Sect. Cycloglottis- Astragalus contortuplicatus

Sect. Swerzowia- Astragalus compositus

Sect. Ankylotus- Astragalus gracilipes, A. affghanus

Sect. Ophiocarpus- Astragalus ophiocarpus

Sect. Harpilobus- Astragalus corrugatus, A. campylorhynchus, A. hauarensis

Sect. Hemiphaca- Astragalus macropterus, A. densiflorus, A. melanostachys, A. clarkeanus

Sect. Christiana- Astragalus orthocarpus, A. zarghumensis, A. tephrosoides

Sect. Chlorostachys- Astragalus chlorostachys, A. bakeri, A. stewarti, A. concretus, A. maxwellii

Sect. Diplotheca- Astragalus graveolens

Sect. Heterozyx- Astragalus neubauerianus

Sect. Cenantrum- Astragalus frigidus, A. pseudochlorostachys

Sect. Himiphragmium- Astragalus oxyodon, A. maddenianus, A. himalayanus, A. strictus A. confertus

Sect. Lithophilus- Astragalus munroi, A. kashmirensis

Sect. Scheremeteviana- Astragalus scheremeteviana

Sect. Komaroviella- Astragalus jabbor-khailii, A. lowarensis

Sect. Malacothrix- Astragalus pseudo-beckii

Sect. Erionotus- Astragalus orbiclautus, A. chitralensis, A. polybotrys

Sect.Myobroma- Astragalus fernandezianus, A. concinnus, A. toppinianus, A. flexus, A. auganus, A. rhizocephalus, A. candolleanus, A. minutofoliolatus, A. gompholobium A. pyrrhotrichus, A. malacophyllus, A. webbianus, A. flemingii, A. staintonianus, A. rhizanthus, A. zanskaresis, A. pindreensis, A. nicharensis, A. purpurascens, A. gilgitensis, A. edelbergianus, A. sultani, A. sp. A

Sect. Cartillaginella- Astragalus retamocarpus

Sect. Chronopus- Astragalus albo-villosus

Sect. Mirae- Astragalus mirus


1 Vexillum pilose externally   (2)
+ Vexillum glabrous   (5)
       
2 (1) Plant 8 cm or less long; calyx longer than the vexillum, vexillum 8.5 mm or less long   Astragalus mirus
+ Plant more than 10 cm long; calyx shorter than the vexillum, vexillum more than 10 mm long   (3)
       
3 (2) Stipules longer than the internodes (aerial stem almost absent)   (4)
+ Stipules shorter than the internodes   Astragalus orbiculatus
       
4 (3) Vexillum more than 3 cm long   Astragalus fernandezianus
+ Vexillum less than 2 cm long   Astragalus chitralensis
       
5 (1) Leaflets verticillate   (6)
+ Leaflets not verticillate   (8)
       
6 (5) Calyx covered with spreading hairs   Astragalus neubauerianus
+ Calyx glabrous or only minutely hairy in the upper portion   (7)
       
7 (6) Raches less than 15 cm long, leaflets 3-6 in each whorl, c. 1-2 mm broad   Astragalus toppinianus
+ Raches more than 15 cm long, leaflets 4-12 in each whorl, c. 2-3 mm broad   Astragalus sp._A
       
8 (5) Flowers sessile or subsessile, forming a compact head   (9)
+ Flowers pedicellate, not forming a compact head   (16)
       
9 (8) Fruit curved at the base   (10)
+ Fruit not curved at the base   (12)
       
10 (9) Fruit 12 mm or less long, glabrous or with appressed hairs   (11)
+ Fruit more than 13 mm long, with appressed and spreading hairs   Astragalus schimperi
       
11 (10) Fruit with appressed hairs   Astragalus tribuloides var. tribuloides
+ Fruit glabrous to subglabrous (with a few hairs at the tip)   Astragalus tribuloides var. leiocarpus
       
12 (9) Fruit as long as the calyx, unilocular   Astragalus leucocephalus
+ Fruit longer than the calyx, semibilocular-bilocular   (13)
       
13 (12) Calyx cup 1-2 mm long, lower teeth x. 4 mm long, upper teeth c. 2.5 mm long, fruit rugose (Illustration.)   Astragalus contortuplicatus
+ Calyx cup c. 2-2.5 mm long, teeth c. 2 mm or less long   (14)
       
14 (13) Leaflets emarginate, fruit ovate   (15)
+ Leaflets not emarginate, obtuse, rarely retuse; fruit semilunar, arcuate   Astragalus filicaulis
       
15 (14) Fruit glabrous (Illustration.)   Astragalus ammophilus var. ammophilus
+ Fruit pubescent (Illustration.)   Astragalus ammophilus var. persepolitanus
       
16 (8) Calyx totally glabrous (keel longer than the wing and the vexillum)   Astragalus concinnus
+ Calyx not totally glabrous   (17)
       
17 (16) Fruit sessile or stipe less than 1 mm long   (18)
+ Fruit stipitate, stipe more 1 mm long   (62)
       
18 (17) Internodes longer than the stipules   (19)
+ Internodes shorter than the stipules   (45)
       
19 (18) Fruit abruptly curved near the tip   (20)
+ Fruit not abruptly curved near the tip   (23)
       
20 (19) Vexillum appendiculate (Illustration). (fruit 1.5-1.7 cm long, septum extending up to half the length of the fruit). (Illustration.)   Astragalus gracilipes
+ Vexillum not appendiculate   (21)
       
21 (20) Fruit long, slender, 1-2 mm broad, slightly torulose   Astragalus ophiocarpus
+ Fruit more than 2 mm broad, not torulose   (22)
       
22 (21) Fruit glabrous, corrugated (Illustration.)   Astragalus corrugatus
+ Fruit pubescent, not corrugated   Astragalus campylorrhynchus
       
23 (19) Margin of the fruit spiny (Illustration.)   Astragalus compositus
+ Margin of the fruit not spiny   (24)
       
24 (23) Fruit glabrous or glabrescent   (25)
+ Fruit pilose   (28)
       
25 (24) Leaf raches hard, spiny   Astragalus webbianus
+ Leaf raches not hard and spiny   (26)
       
26 (25) Plant covered with white and black spreading hairs, leaflets more than 15 pairs (Illustration.)   Astragalus retamocarpus
+ Plant glabrous or with white appressed hairs only, leaflets 8 pairs or less   (27)
       
27 (26) Inflorescence elongated, more than 5 cm long with more than 10 racemosely arranged flowers; fruit longitudinally dorsally grooved   Astragalus macropterus
+ Inflorescence less than 5 cm long, with 1-3, 3-6-flowered clusters, fruit without grooves (Illustration.)   Astragalus oxyglottis
       
28 (24) Stipules almost encircling the stem (at least more than half the girth)   (29)
+ Stipules not encircling the stem   (36)
       
29 (28) Plant (except the old stem) villous, calyx cup more than 5 mm (inflorescence lax.)   Astragalus polybotrys
+ Plant not villous, calyx cup less than 5 mm   (30)
       
30 (29) Bracts longer than the calyx, fruit 2-seeded   Astragalus melanostachys
+ Bracts shorter than the calyx, fruit more than 2-seeded   (31)
       
31 (30) Fruit arcuate, semilunar   (32)
+ Fruit not as above   (35)
       
32 (31) Calyx teeth shorter than the cup   (33)
+ Calyx teeth longer than the cup   (34)
       
33 (32) Keel 5-7 mm long   Astragalus strictus
+ Keel 8-10 mm long   Astragalus confertus
       
34 (32) Wing longer than or subequal to the keel; leaflets c. 2-3.5 mm long, c.1 mm broad; vexillum appreciably longer than the keel   Astragalus jabbor-khailii
+ Wing shorter than the keel; leaflets 3.0-5.0 mm long, 1.5-2.5 mm broad; vexillum subequal or slightly longer than the keel   Astragalus lowarensis
       
35 (31) Fruit rugose (Illustration.)   Astragalus densiflorus
+ Fruit not rugose   Astragalus clarkeanus
       
36 (28) Inflorescence a terminal panicle, 25 cm or more long   Astragalus tephrosoides var. lacei
+ Inflorescence not a terminal panicle, not as long as above   (37)
       
37 (36) Flower less than 1 cm long (c. 6-7 mm), fruit arcuate   (38)
+ Flower more than 1 cm long   (39)
       
38 (37) Fruit more than 12 (c. 20)-seeded, more than 12 mm long   Astragalus hauarensis var. hauarensis
+ Fruit 5-12-seeded, c. 12 mm long   Astragalus hauarensis var. brachycarpus
       
39 (37) Calyx teeth equal to or longer than the cup   (40)
+ Calyx teeth shorter than the cup   (43)
       
40 (39) Calyx cylindrical, keel c. 2.5 mm broad   Astragalus orthocarpus
+ Calyx campanulate, keel c. 4.5 mm broad   (41)
       
41 (40) Leaflets 31-37 per leaf, c. 5-10 mm long   Astragalus zarghumensis
+ Leaflets 49-59 per leaf, 8-30 mm long   (42)
       
42 (41) Plant villous, hairs white   Astragalus albovillosus var. albovillosus
+ Plant villous, hairs black and white   Astragalus albovillosus var. nigrescens
       
43 (39) Petiole less than 1 cm long; calyx teeth c. 5 mm long   Astragalus polybotrys
+ Petiole more than 1 cm long   (44)
       
44 (43) Leaflets c. 4-10 mm long, c. 2-5 mm broad; peduncle 8-50 mm long   Astragalus malacophyllus
+ Leaflets 11-20 mm long, c. 5-12 mm broad; peduncle 4.5-11.5 mm long   Astragalus kashmirensis
       
45 (18) Fruit curved inward abruptly near the tip (vexillum appendiculate)   (46)
+ Fruit not as above   (47)
       
46 (45) Fruit 1.5-1.7 cm long, septum extending up to half the length of the fruit (Illustration.)   Astragalus gracilipes
+ Fruit 2-3.5 cm long, septum extending more than half the length of the fruit (Illustration.)   Astragalus affghanus
       
47 (45) Ovary and fruit glabrous   (48)
+ Ovary and fruit pilose   (49)
       
48 (47) Leaflets less than 3 mm long   Astragalus minuto-foliolatus
+ Leaflets more than 3 mm long   Astragalus webbianus
       
49 (47) Flower less than 1 cm (c. 6-9 mm) long, fruit arcuate   (50)
+ Flower more than 1 cm long, fruit not as above   (52)
       
50 (49) Leaflets c. 1 mm broad, fruit less than 1 cm long   Astragalus confertus
+ Leaflets more than 1 mm broad, fruit more than 1 cm long   (51)
       
51 (50) Fruit 12(-20)-seeded, more than 12 mm long   Astragalus hauarensis var. hauarensis
+ Fruit 5-12-seeded, c. 12 mm long   Astragalus hauarensis var. brachycarpus
       
52 (49) Leaflets acute   (53)
+ Leaflets not acute   (54)
       
53 (52) Leaf c. 6 cm long, leaflets velvety tomentose on both sides   Astragalus sultani
+ Leaf more than 10 cm long, leaflets glabrous above, pilose below   Astragalus gompholobium
       
54 (52) Leaflets orbicular to suborbicular   Astragalus pyrrhotrichus
+ Leaflets not as above   (55)
       
55 (54) Peduncle 6-31.5 cm long (Illustration.)   Astragalus flemingii
+ Peduncle less than 5 cm long   (56)
       
56 (55) Lower calyx teeth 5 mm or more long   (57)
+ Lower calyx teeth 3-4 mm long   (59)
       
57 (56) Inflorescence pedunculate, peduncle 8-50 cm long   Astragalus malacophyllus
+ Inflorescence sessile or peduncle 5 mm long   (58)
       
58 (57) Upper calyx teeth c. 6 mm long, seeds c. 8 per fruit   Astragalus staintonianus
+ Upper calyx teeth c. 3-4 mm long, seeds more than 10 per fruit   Astragalus rhizanthus
       
59 (56) Leaflets 15-25, rarely 31 per leaf   (60)
+ Leaflets 39 or more per leaf   (61)
       
60 (59) Rachis stout, erect   Astragalus zanskarensis
+ Rachis not stout, curved   Astragalus pindreensis
       
61 (59) The distance between the two successive pairs of leaflets more than 3 mm   Astragalus nicharensis
+ The distance between the successive pairs of leaflets c. 3 mm or less (Illustration.)   Astragalus purpurascens
       
62 (17) Internodes longer than the stipules   (63)
+ Internodes shorter than the stipules   (79)
       
63 (62) Ovary and pod glabrous   (64)
+ Ovary and pod pubescent   (69)
       
64 (63) Calyx teeth deltoid, c. 1.5 mm or less long   (65)
+ Calyx teeth not deltoid, longer than 1.5 mm   (66)
       
65 (64) Leaflets 7 per leaf, elliptic-ovate   Astragalus pseudochlorostachys
+ Leaflets more than 7(13-17) per leaf   Astragalus chlorostachys
       
66 (64) Stipules not foliaceous, c.1 mm broad   (67)
+ Stipules foliaceous, c. 4 mm or more broad   (68)
       
67 (66) Fruit 10-12-seeded, c. 1.5-1.8 cm long (excluding the stipe and the style) (Illustration.)   Astragalus bakeri
+ Fruit 6-8-seeded , c. 1.3 cm or less long (excluding the stipe and the style) (Illustration.)   Astragalus stewartii
       
68 (66) Leaflets oblong to oblong-lanceolate, fruit curved, not ensiform (Illustration.)   Astragalus concretus
+ Leaflets round elliptical ovate, fruit straight, ensiform (Illustration.)   Astragalus graveolens
       
69 (63) Fruit arcuate   Astragalus strictus
+ Fruit not arcuate   (70)
       
70 (69) Stipules foliaceous, more than 1 cm broad   Astragalus kuramensis
+ Stipules not as above   (71)
       
71 (70) Fruit unilocular   (72)
+ Fruit bilocular (partly or completely)   (75)
       
72 (71) Calyx teeth c. 1 mm or less   Astragalus frigidus
+ Calyx teeth more than 1 mm long   (73)
       
73 (72) Leaflets elliptic, acute   Astragalus oxyodon
+ Leaflets oblong, obtuse-truncate or emarginate   (74)
       
74 (73) Calyx teeth equal to or longer than the cup   Astragalus maddenianus
+ Calyx teeth shorter than the cup (cup c. 3 mm, teeth c. 1-2 mm)   Astragalus himalayanus
       
75 (71) Fruit oblong, gibbous, whole plant covered with dense white spreading hairs   Astragalus munroi
+ Not as above   (76)
       
76 (75) Stipe shorter than the calyx cup   (77)
+ Stipe longer than the calyx cup   Astragalus maxwellii
       
77 (76) Calyx teeth c. 1-3 mm long, with black hairs   Astragalus tibetanus
+ Calyx teeth 4 mm or more long, with yellowish or black and white hairs   (78)
       
78 (77) Plant villous, upper calyx teeth 4 mm, lower 5 mm long, with black and white hairs. (Illustration.)   Astragalus pseudo-beckii
+ Plant not villous, upper calyx teeth 5 mm, lower 6 mm long, with yellowish hairs   Astragalus candolleanus
       
79 (62) Ovary and fruit glabrous   (80)
+ Ovary and fruit pubescent   (81)
       
80 (79) Leaflets and calyx subpubescent   Astragalus edelbergianus
+ Leaflets and calyx uniformly pubescent   Astragalus sechremetevianus
       
81 (79) Leaflets orbicular-ovate or orbicular-obovate   Astragalus flexus
+ Leaflets not orbicular   (82)
       
82 (81) Fruit 11-12 mm broad. (Illustration.)   Astragalus auganus
+ Fruit c. 6 mm or less broad   (83)
       
83 (82) Vexillum panduriform   Astragalus rhizocephalus
+ Vexillum not panduriform   (84)
       
84 (83) Calyx teeth 9-10 mm long, longer than the cup   Astragalus gilgitensis
+ Calyx teeth 6-6.5 mm long, shorter than the cup   Astragalus candolleanus

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