1 |
Spike single, terminal |
|
(2) |
+ |
Spikes 2 to numerous, usually 1 to several spikes borne in an involucral bract sheath, rarely arranged in compound inflorescence |
|
(8) |
|
|
|
|
2 (1) |
Nutlets biconvex or plano-convex, stigmas 2 |
|
(3) |
+ |
Nutlets trigonous, stigmas 3 |
|
(4) |
|
|
|
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3 (2) |
Spikes bisexual, androgynous; utricles not spongy at base. |
|
51 C. sect. Capituligerae |
+ |
Spikes unisexual; utricles spongy at base. |
|
52 C. sect. Physoglochin |
|
|
|
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4 (2) |
Female glumes early deciduous; utricles downward reflexed in maturity, lateral veins not more conspicuous than veins on abaxial surface; regressive branch rachilla conspicuous, usually longer than nutlets. |
|
46 C. sect. Leucoglochin |
+ |
Female glumes deciduous at same time as utricles or later than utricles; utricles not distinctly downward reflexed in maturity, lateral veins more conspicuous than veins on abaxial surface; without regressive branch rachilla, or if with regressive branch rachilla then rachilla shorter than nutlets |
|
(5) |
|
|
|
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5 (4) |
Spikes shortly cylindric to capitate |
|
(6) |
+ |
Spikes linear or linear-cylindric, rarely only female spikes so |
|
(7) |
|
|
|
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6 (5) |
Nutlets with regressive branch rachilla at base. |
|
47 C. sect. Rupestres |
+ |
Nutlets without regressive branch rachilla at base. |
|
48 C. sect. Rarae |
|
|
|
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7 (5) |
Utricles shortly beaked (beak not over 1 mm), veins obscure on abaxial surface. |
|
49 C. sect. Grallatoriae |
+ |
Utricles long beaked (beak over 1 mm), veins conspicuous on abaxial surface. |
|
50 C. sect. Rhizopodae |
|
|
|
|
8 (1) |
Utricles plano-convex or biconvex; stigmas 2 |
|
(9) |
+ |
Utricles trigonous; stigmas 3 |
|
(13) |
|
|
|
|
9 (8) |
Involucral bracts sheathed; female spikes sparsely flowered, rarely slightly densely flowered. |
|
45 C. sect. Graciles |
+ |
Involucral bracts without sheaths; female spikes densely many flowered |
|
(10) |
|
|
|
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10 (9) |
Spikes pedunculate, ± pendent |
|
(11) |
+ |
Spikes nearly sessile or only lowermost or basal spike rather long pedunculate |
|
(12) |
|
|
|
|
11 (10) |
Spikes 2-6 borne in every involucral bract sheath on inflorescence. |
|
44 C. sect. Tuminenses |
+ |
Spikes solitary in every involucral bract sheath on inflorescence. |
|
43 C. sect. Praelongae |
|
|
|
|
12 (10) |
Utricles distinctly beaked, beak 0.5-1.5 mm, 2-toothed at orifice. |
|
42 C. sect. Forficulae |
+ |
Utricles beakless or shortly beaked, truncate or emarginate at orifice. |
|
41 C. sect. Phacocystis |
|
|
|
|
13 (8) |
Utricles nearly beakless or with short beak, truncate, emarginate, or very shortly 2-toothed at orifice |
|
(14) |
+ |
Utricles with a long beak or medium-sized beak (beak rather short in C. sect. Paludosae), long or shortly 2-toothed at orifice, rarely subtruncate or very shortly toothed |
|
(28) |
|
|
|
|
14 (13) |
Female spikes densely flowered; involucral bracts usually without sheath or only very shortly sheathed |
|
(15) |
+ |
Female spikes sparsely flowered; involucral bracts usually sheathed |
|
(20) |
|
|
|
|
15 (14) |
Female spikes remote, pedunculate |
|
(16) |
+ |
Female spikes contiguous or rather congregate, rarely lower 1 or 2 remote, without peduncle or very shortly pedunculate, rarely lowermost one rather long pedunculate |
|
(17) |
|
|
|
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16 (15) |
Male spikes 2-6 (only 1 male spike in C. inanis); utricles hispidulous or scabrous. |
|
15 C. sect. Hirtae |
+ |
Male spike solitary, terminal; utricles glabrous. |
|
16 C. sect. Anomalae |
|
|
|
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17 (15) |
Involucral bracts leaflike or lower ones leaflike, rarely setaceous; terminal spike gynaecandrous, rarely male. |
|
11 C. sect. Racemosae |
+ |
Involucral bracts scale-shaped or lower ones linear, upper ones scale-shaped; terminal spike male |
|
(18) |
|
|
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18 (17) |
Female glumes red-brown or dark brown; utricles glabrous, usually shiny, obliquely truncate at orifice of beak; style base thickened. |
|
14 C. sect. Lamprochlaenae |
+ |
Female glumes brown or brownish; utricle hirtellous or papillate, emarginate or very shortly 2-toothed at orifice of beak |
|
(19) |
|
|
|
|
19 (18) |
Utricles papillate; style base thickened. |
|
12 C. sect. Glauciformes |
+ |
Utricles hirtellous; style base not thickened. |
|
13 C. sect. Acrocystis |
|
|
|
|
20 (14) |
Utricles glabrous or scabrid |
|
(21) |
+ |
Utricles pubescent or hispidulous, rarely glabrous |
|
(24) |
|
|
|
|
21 (20) |
Leaf blades usually revolute at margins; involucral bracts sheathlike, without blades; male spike few flowered; style base thickened and globose. |
|
22 C. sect. Albae |
+ |
Leaf blades flat; involucral bracts leaflike or spathelike, with short blades; male spike many flowered; style base not thickened |
|
(22) |
|
|
|
|
22 (21) |
Culms lateral; leaf blades rather broad, 6-30 mm wide (very few species with 3-6 mm wide blades); spikes bisexual, androgynous; female glumes pale or reddish brown. |
|
25 C. sect. Siderostictae |
+ |
Culms central; leaf blades rather narrow, 2-5 mm wide; spikes unisexual; female glumes ferruginous-brown or yellow-brown |
|
(23) |
|
|
|
|
23 (22) |
Involucral bracts shortly sheathed; female spikes densely flowered, slenderly pedunculate, usually pendulous; style exserted from utricle. |
|
23 C. sect. Limosae |
+ |
Involucral bracts long sheathed; female spikes sparsely flowered, slightly thickly pedunculate, erect; style included in utricle (only stigmas exserted). |
|
24 C. sect. Paniceae |
|
|
|
|
24 (20) |
Involucral bracts spathelike, without blade, rarely leaflike; nutlets not thickened at apex into annulate shape; style base slightly thickened or not thickened, rarely thickened and conic |
|
(25) |
+ |
Involucral bracts with short blades; nutlets thickened at apex into annulate shape; style base thickened into different shapes |
|
(26) |
|
|
|
|
25 (24) |
Involucral bracts leaflike, rarely sheathlike; spikes bisexual, androgynous; utricles subleathery or papery; style base thickened and conic. |
|
20 C. sect. Radicales |
+ |
Involucral bracts spathelike, rarely sheathlike; spikes unisexual, terminal spike male, lateral spikes female; utricles membranous; style base thickened or not thickened. |
|
21 C. sect. Clandestinae |
|
|
|
|
26 (24) |
Nutlets with persistent and bent style base at apex. |
|
17 C. sect. Cryptostachyae |
+ |
Nutlets without persistent style base at apex, or if nutlets with persistent style base then style base straight or thickened into cucullate shape |
|
(27) |
|
|
|
|
27 (26) |
Nutlets with ± cylindric style base, concave at apex, persistent style base not thickened into cucullate shape. |
|
18 C. sect. Lageniformes |
+ |
Nutlets ± thickened into disk shape, not concave at apex, persistent style base thickened into cucullate shape. |
|
19 C. sect. Mitratae |
|
|
|
|
28 (13) |
Leaf blades with transverse septate nodes between longitudinal veins |
|
(29) |
+ |
Leaf blades without transverse septate nodes between longitudinal veins |
|
(33) |
|
|
|
|
29 (28) |
Utricles ± hispidulous or pubescent, rarely glabrous or ciliate on margins of beak |
|
(30) |
+ |
Utricles glabrous |
|
(31) |
|
|
|
|
30 (29) |
Female glumes ovate or broadly ovate, apex acute, mucronate; utricles obtusely trigonous, ca. 4 mm, rarely 5 mm, shortly beaked, shortly 2-toothed at orifice. |
|
39 C. sect. Occlusae |
+ |
Female glumes lanceolate, narrowly ovate, or oblong-ovate (broadly ovate in C. latisquamea), apex acuminate, aristate or mucronate; utricles slightly inflated trigonous or inflated trigonous, 5-10 mm, rarely ca. 4 mm, medium to long beaked, ± long to long 2-toothed at orifice. |
|
40 C. sect. Carex |
|
|
|
|
31 (29) |
Female glumes long aristate; utricles eventually horizontally patent or downward reflexed, obtusely trigonous, long beaked, deeply splitting into 2 long teeth at orifice. |
|
36 C. sect. Pseudocypereae |
+ |
Female glumes muticous or mucronate, rarely aristate; utricles eventually obliquely patent, rarely horizontally patent or downward reflexed, inflated trigonous or slightly so, medium to long beaked or very shortly beaked |
|
(32) |
|
|
|
|
32 (31) |
Female glumes oblong-lanceolate or narrowly ovate, without mucro or awn; utricles inflated trigonous, membranous or subleathery, medium to long beaked. |
|
37 C. sect. Physocarpae |
+ |
Female glumes ovate or broadly ovate, mucronate, rarely aristate; utricles slightly inflated trigonous, leathery or suberose, very shortly beaked. |
|
38 C. sect. Paludosae |
|
|
|
|
33 (28) |
Female glumes dark purple-red or brown; utricles compressed trigonous, rarely trigonous |
|
(34) |
+ |
Female glumes pale, rarely dark brown; utricles trigonous or inflated trigonous (subplano-convex in C. sect. Secalinae) |
|
(35) |
|
|
|
|
34 (33) |
Spikes numerous, bisexual, androgynous, usually several spikes borne in an involucral bract sheath. |
|
26 C. sect. Decorae |
+ |
Spikes 2-7, unisexual, solitary in an involucral bract sheath, terminal spikes male, rarely gynaecandrous, lateral spikes female. |
|
27 C. sect. Aulocystis |
|
|
|
|
35 (33) |
Culms lateral, rarely central; nutlets usually constricted or with constriction on angles, rarely not constricted or without constriction; style base thickened or inflated, rarely not thickened |
|
(36) |
+ |
Culms central; nutlets not constricted or without constrictions on angles; style base not thickened |
|
(37) |
|
|
|
|
36 (35) |
Female spikes densely many flowered; nutlets usually constricted or with constriction on angles, rarely not constricted. |
|
34 C. sect. Rhomboidales |
+ |
Female spikes sparsely several flowered; nutlets not constricted on angles. |
|
35 C. sect. Careyanae |
|
|
|
|
37 (35) |
Male spikes 1-3, borne at apex of culm; utricles leathery, subplano-convex. |
|
33 C. sect. Secalinae |
+ |
Terminal spike male, rarely androgynous or gynaecandrous; utricles membranous or papery, rarely subleathery |
|
(38) |
|
|
|
|
38 (37) |
Female spikes densely many flowered; involucral bracts usually without sheaths or with very short sheath; utricles eventually divaricate, horizontally patent, or downward reflexed, rarely obliquely patent, distinctly several veined |
|
(39) |
+ |
Female spikes usually sparsely flowered (densely flowered in C. sect. Hymenochlaenae); involucral bracts sheathed; utricles erect or slightly obliquely patent, greenish or greenish yellow, obscurely veined |
|
(40) |
|
|
|
|
39 (38) |
Female glumes long aristate, rarely mucronate (C. ischnostachya and C. subtumida without mucro or awn); utricles eventually horizontally patent or slightly downward reflexed, rarely slightly divaricate, brown- green or dark brown or black-brown at maturity. |
|
31 C. sect. Confertiflorae |
+ |
Female glumes mucronate or muticous, rarely aristate; utricles eventually obliquely patent, rarely horizontally patent, yellowish green or stramineous, rarely brownish yellow. |
|
32 C. sect. Molliculae |
|
|
|
|
40 (38) |
Female spikes oblong or shortly cylindric; utricles less than 3 mm. |
|
28 C. sect. Chlorostachyae |
+ |
Female spikes cylindric, narrowly cylindric, or linear- cylindric; utricles 4-7 mm, rarely 3-3.5 mm |
|
(41) |
|
|
|
|
41 (40) |
Female spikes cylindric, rather densely flowered; utricles elliptic or obovate- elliptic, shortly 2-toothed at orifice. |
|
29 C. sect. Hymenochlaenae |
+ |
Female spikes narrowly cylindric or linear- cylindric, sparsely or very loosely flowered; utricles oblong-lanceolate, lanceolate, or narrowly oblong, obliquely truncate or very shortly 2-toothed at orifice. |
|
30 C. sect. Debiles |
|